Bark and wood anatomy of the Tribe Hamelieae (Rubiaceae)

TL;DRAbstract

We studied wood and bark anatomy of six ( Deppea, Hamelia, Hoffmannia, Omiltemia, Pinarophyllon , and Plocaniophyllon ) of the seven genera of the tribe Hamelieae sensu Robbrecht, and Syringantha with as main purposes to determine if there are characters that support the boundaries of the Hamelieae, to evaluate the status of Syringantha as a member of the Hamelieae, and to evaluate the taxonomic position of Hamelieae within the subfamilies Rubioideae or Cinchonoideae. In addition, we studied for comparative purposes representative species of Psychotria (Psychotrieae, Rubioideae), Exostema , and Hintonia ( Portlandia group, Cinchonoideae), Randia (Gardenieae, Ixoroideae), and Bouvardia ( incertae sedis ). Bark of most genera studied had a single periderm, while a rhytidome was observed in Exostema and few species of Psychotria . The mineral inclusions allowed recognizing related genera, for example, raphides in Hamelieae and Psychotria , prisms in Exostema , and druses in Randia. Member

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